Quinn's DC Universe comic book origin, revealed in Batman: Harley Quinn (October 1999), is largely an adaptation of her animated origin from the Mad Love graphic novel. A Harley Quinn ongoing series was published monthly by DC Comics for 38 issues from 2001 to 2003. Creators who contributed to the title included Karl Kesel, Terry Dodson, A.J. Lieberman, and Mike Huddleston. The series dealt with her going solo, eventually starting a gang and then fleeing Gotham for the city of Metropolis with her friend Poison Ivy. Quinn dies, only to be resurrected and return to Gotham. The series ends with Harley turning herself in to Arkham Asylum, having finally understood that she needs help. We also learn in issue #8 of the comic that Harley had a relationship in college with fellow psychiatry major Guy Kopski whose suicide started her obsession with the Joker. Harley later appears in the Jeph Loeb series Hush. She is next seen in a Villains United Infinite Crisis special, where she is one of the many villains who escape from Arkham (although she is knocked unconscious the moment she escapes).
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